RALPH: Pigeons, like doves, are docile, tame birds; while gall is secreted by the liver was thought to be the source of anger. Hamlet accuses himself of lacking gall, unable to get angry enough to find the taste of oppression bitter.

SARAH: If he could feel that anger, he would have fed Claudius's guts to the vultures by now.

RALPH: The last few lines here are directed towards Claudius. It's as if thinking about Claudius's crimes has called up his image up in front of Hamlet, and finally gotten him angry — so now these insults are finally not directed at himself, but at his enemy.